FVSUag Directory


Nirmal Joshee, PhD

Professor of Plant Science

(478) 822-7039

(478) 825-6373


Academic Program: Biotechnology Graduate Program

Research Area: Bioenergy and Climate Change, Plant Biotechnology

Research Interests:
  • Medicinal plants and trichome biology
  • Plant tissue culture and genetic transformation
  • Reproductive biology
  • Plants and value added products

3 photos of various research subjects including plants and a scanning electron micrograph

Dr. Joshee and colleagues pose for a photo while in a laboratory on the campus of Fort Valley State University


Doctor of Philosophy: Botany - North Eastern Hill University, Meghalaya, India (1987)

Master of Science: Botany - Kumaun University, Naini Tal, Uttarakhand, India (1980)

Bachelor of Science: Biology - Kumaun University, Naini Tal, Uttarakhand, India (1978)

Recent Publications/Presentations:

Books Edited

Medicinal Plants: Farm to Pharmacy. Springer. 2019.

Nirmal Joshee, S. Dhekney, and P. Parajuli (Editors). ISBN 978-3-030-31268-8

Book Chapters/Proceedings

Shahin, L, Mahapatra, AK, Joshee N. 2019. Effect of various drying methods on drying characteristics and antioxidant properties of Paulownia elongata leaves. IGA Asian Regional Conference on Goats (ARCG 2019). Rampur, Chitwan, Nepal, October 20-23. (Submitted for proceedings).

L Irvin, C Jackson, Aisha L. Hill, R Bajaj, C Mahmoudi, Brajesh N. Vaidya, and N Joshee. 2019. Skullcaps (Scutellaria spp.): Ethnobotany and current research. Ch. 7. In: Medicinal Plants: From Farm to Pharmacy. N. Joshee, S. A. Dhekney, P. Parajuli (Editors). Springer. Pp. 141-168.

Robert E. Wright III, Nirmal Joshee, and Prahlad Parajuli. 2019. Modulation of Tumor Immunity by Medicinal Plant or Functional Food-Derived Compounds. Ch. 12. In: Medicinal Plants: From Farm to Pharmacy. N. Joshee, S. A. Dhekney, P. Parajuli (Editors). Springer. Pp. 275-290.

Hussain F, Mittal S, Joshee N and Parajuli P. 2016. Application of Bioactive Compounds from Scutellaria in Neurologic Disorders. In: The Benefits of Natural Products for Neurodegenerative Diseases, Mohamed M, Mohamed A and Gullemin G (Eds), Springer-Verlag, Heidelberg Germany, Chapter 5.

Peer-Reviewed Papers


Shahin, L, SS. Phaal, Brajesh N Vaidya, Jame E. Brown, N Joshee. 2019. Aronia (Chokeberry): an underutilized, highly nutraceutical plant. J Medicinally Active Plants 8(4):46-63.

Yi W, N Joshee, W Cao, Q Wu, and N Tahir. 2019. Continuous hydrogen production by dark and photo co-fermentation using a tubular multi-cycle bio-reactor with Paulownia biomass. Cellulose. 1-10. https://doi.org/10.1007/s10570-019-02468-z.(0123456789().,)0123456789().,-)

Liu Bing, Yun-Hong Tan, Liu Su, Olmstead Richard, Min Dao-Zhang, Chen Zhi-Duan, Joshee N, Vaidya Brajesh, Chung Richard, Li Bo. 2019. Phylogenetic relationships of Cyrtandromoea and Wightia revisited: a new tribe in Phrymaceae and a new family in Lamiales. J. of Systematics and Evolution, 58(1):1-17, May 2019 https://doi.org/10.1111/jse.12513.

Vaidya, B.N., B. Asanakunov, L. Shahin, H.L. Jernigan, N. Joshee and S.A. Dhekney. 2019. Improving micropropagation of Mentha x piperita L. using a liquid culture system. In Vitro Cell & Dev Biology – Plant. 55(1):71–80. https://doi.org/10.1007/s11627-018-09952-4


Soundappan K, Bajaj R, Vaidya BN, Joshee N. 2018. Anatomical and biochemical investigations on medicinal tree Murraya koenigii Spreng. Journal of Biotech Research 9:69-77.

Ramdoss, N, Gupta, D., Vaidya, BN, Joshee, N, Basu, C. 2018. Functional characterization of 1-aminocyclopropane-1-carboxylic acid oxidase gene in Arabidopsis thaliana and its potential in providing flood tolerance. Biochemical and Biophysical Research Communications 503:365-370.

Stewart, WM, Vaidya, BN, Mahapatra AK, Terrill, TH, and Joshee, N. 2018. Potential use of multipurpose Paulownia elongata tree as an animal feed resource. American J Plant Sci. 9:1212-1227.

Tisserat BH., HongSik Hwang, Steven F. Vaughn, Mark A. Berhow, Steven C. Peterson, N Joshee, Brajesh N. Vaidya, and O-kuru RH. 2018. Fiberboard created using the natural adhesive properties of distillers dried grains with solubles. BioResources 13(2):2678-2701.

Peterson, SC and Joshee, N. 2018. Co-milled silica and coppiced wood biochars improve elongation and toughness in styrene-butadiene elastomeric composites while replacing carbon black. Journal of Elastomers and Plastics, 1-10, DOI: 10.1177/0095244317753653


Chen, H, Liu, N, Qu, X, Joshee, N, Liu, S. 2017. The effect of hot water pretreatment on the heavy metal adsorption capacity of acid insoluble lignin from Paulownia elongata. Journal of Chemical Technology & Biotechnology 93(4): 1105-1112. DOI 10.1002/jctb.5469

Chaires, M, Gupta, D, Joshee, N, Cooper, Kerry K, and Basu, C. 2017. RNA-seq analysis of the salt stress induced transcripts in fast growing bioenergy tree, Paulownia elongata. J Plant Interaction 12(1):128–136.

Vaughn SF, Kenar JA, Tisserat B, Jackson MA., N Joshee, Vaidya BN, and Peterson, SC. 2017. Chemical and physical properties of Paulownia elongata biochar modified with oxidants for horticultural applications. Industrial Crops and Products 97:260-267.


Fira, A, N Joshee, V Cristea, M Simu, M Harta, D Pamfil, D Clapa. 2016. Optimization of micropropagation protocol for Goji berry (Lycium barbarum L.). Bulletin UASVM Horticulture 73(2):142-150.

Yan, J, Joshee, N, and Liu, S. 2016. Utilization of hardwood in biorefinery: A kinetic interpretation of pilot-scale hot-water pretreatment of Paulownia elongata woodchips. J. Biobased Materials and Bioenergy 10:339-348.

Vaidya BN, CL Jackson, ZD. Perry, SA Dhekney, and N Joshee. 2016. Agrobacterium-mediated transformation of thin cell layer explants of Scutellaria ocmulgee Small: A rare plant with anti-tumor properties. Plant Cell, Tissue, and Organ Culture, 127(1):57–69; doi:10.1007/s11240-016-1029-y.

Croom, AL, C. Jackson, BN Vaidya, P Parajuli, and N Joshee. 2016. Thin cell layer (TCL) culture system for herbal biomass production and genetic transformation of Bacopa monnieri L. Wettst. American J Plant Sciences 7: 1232-1245.

Ting He, BN Vaidya, ZD. Perry, P Parajuli, N Joshee. 2016. Paulownia as a Medicinal Tree: Traditional Uses and Current Advances. European J Medicinal Plants 14(1):1-15.

A Similien, DA Shannon, C. Wesley Wood, AM Rimando, BW Kemppainen, Edzard van Santen and N Joshee. 2016. Shade, irrigation and nutrients affect flavonoid concentration and yield in american skullcap. Crop Science, 56(3):1213-1224. doi:10.2135/cropsci2015.03.0186


Vaughn SF,  F. Dan Dinelli, Brent Tisserat, Nirmal Joshee, Martha M. Vaughan, Steven C. Peterson. 2015. Creeping bentgrass growth in sand-based root zones with or without biochar. Scientia Horticulturae 197: 592-596.

Basu C, N. Joshee, T. Gazelian, B. N. Vaidya, A. Saditkit, H. Hemmati, and Z. D Perry. 2015. Cross-species PCR and field studies on Paulownia elongata: A potential bioenergy crop. Bioethanol 2:12-23.

Shiwakoti S, D. A. Shannon, C. W. Wood, N. Joshee, A. Rimando, K. S. Lawrence and B. Kemppainen. 2015. Nitrogen, phosphorus and potassium effects on biomass yield and flavonoid content of American skullcap (Scutellaria lateriflora). J. Plant Nutrition. 39(9):1240-1249. DOI: 10.1080/01904167.2015.1050509

Tisserat B, L, Reifschneider, N Joshee, and V. L. Finkenstadt. 2015. Evaluation of Paulownia elongata Wood Polyethylene Composites. Journal of Thermoplastic Composite Materials. 28:1301-1320. DOI: 10.1177/0892705713505856. First published on September 26, 2013.


Brearley, TA, Brajesh N Vaidya, and N Joshee. 2014. Cytokinin, Carbon Source, and Acclimatization Requirements for In Vitro Propagation of Scutellaria barbata D. Don and Scutellaria racemosa Pers. American Journal of Plant Sciences, 5: 3662-3672.

Marsh Z, Yang, T., Nopo-Olazabal, L., Wu, S., Ingle, T., Joshee, N., and Medina-Bolivar, F. 2014. Effect of light, methyl jasmonate and cyclodextrin on production of polyphenolic compounds in hairy root cultures of Scutellaria lateriflora. Phytochemistry, 107:50-60. http://dx.doi.org/10.1016/j.phytochem.2014.08.020